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D to be catalytically active to transmit stress signals to the Spc1 MAPK. Lys-1066 in the ATP-binding site of Wis4 was substituted with methionine in the wis4:myc strain to create the wis4KM:myc strain, which expresses catalytically inactive Wis4KM-myc at a level comparable to active Wis4-myc (Figure 3A, bottom). Spc1 activation upon osmostress was only slightly reduced in the wis4KM strain and wa

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